Редовен професор д-р Златко Левков



Research Assistant at FNS, Skopje                                      November 1996 - May 2001

Teaching Assistant at FNS, Skopje                                      May 2001- February 2006

Assistant Professor at FNS, Skopje                                     February 2006 - November 2010

Associate Professor at FNS, Skopje                                     November 2010 - November 2015

Full Professor at FNS, Skopje                                              November 2015-



Full Professor at the Faculty of Natural Sciences



Head of Institute of Biology, Faculty of Natural Sciences (2009-2011)

Vice-dean for education at the Faculty of Natural Sciences (2011-2012)


TEACHING SUBJECTS: (graduate and postgraduate level)

  1. Systematic and phylogeny of algae;
  2. General phycology
  3. Ecological monitoring and bioindicators
  4. Forensic Botany
  5. Paleoecology



  • Taxonomy and biogeography of diatoms;
  • Systematic and phylogeny of diatoms
  • Species flocks in ancient lakes;
  • Palaeoecology and palaeolimnology
  • Glacial lakes' diatom composition;
  • Diatoms from estuaries and costal waters;
  • Diatom communities for monitoring of the pollution;
  • Diatoms assemblages in forensic sciences.



MSci in Biology (2001): Distribution of heavy metals in River Vardar and their impact on aquatic mosses and diatom flora.

PhD in Biology (2005): Taxonomy and ecology of diatom flora from Lakes Prespa and St. Naum Springs (Lake Ohrid)




  1. LEVKOV, Z., Krstic, S., Metzeltin, D. & Nakov, T. (2007): Diatoms of Lakes Prespa and Ohrid (Macedonia). Iconographia Diatomologica 16: 1–603. A.R.G. Gantner Verlag K.G.
  2. LEVKOV, Z. (2009): Amphora sensu lato. Diatoms of Europe, Diatoms of the European Inland waters and comparable habitats 5: 1–916. A.R.G. Gantner Verlag. K.G.
  3. Cvetkoska, A., Reed, J. & LEVKOV, Z. (2012): Lake Ohrid diatoms as palaeoclimate indicators of climate change during the last glacial-interglacial cycle. Diatom monographs 15: 1–216. A.R.G. Gantner Verlag. K.G.
  4. LEVKOV, Z., Metzeltin, D. & Pavlov, A. (2013): Luticola and Luticolopsis. Diatoms of Europe, Diatoms of the European Inland waters and comparable habitats 7: 1–697. A.R.G. Gantner Verlag K.G.
  5. LEVKOV, Z., Mitić-Kopanja, D. & Reichardt, E. (2016): The genus Gomphonema in the Republic of Macedonia. Diatoms of the European Inland waters and comparable habitats 8: 1–552. Koeltz Scientific Books.
  6. Jovanovska, E. & LEVKOV, Z. (2020): The genus Diploneis in the Republic of Macedonia. Diatoms of the European Inland waters and comparable habitats 9: 527-689. Koeltz Scientific Books.



  1. Krstic, S., LEVKOV, Z. & Stojanovski, P. (1997a): Diatoms in monitoring of River Vardar, Macedonia. Ekologija 32(2): 1–16.
  2. Krstic, S., LEVKOV, Z. & Stojanovski, P. (1997b): Use of algae for monitoring rivers in Macedonia. In: Prygiel J., Whitton B.A. & Bukowska J. (eds.): Use of Algae for Monitoring Rivers 3: 145–153.
  3. LEVKOV, Z. & Krstic, S. (2002): Use of algae for monitoring of heavy metals in the River Vardar, Macedonia. Mediterrenian Marine Science, 3(1): 101–114.
  4. Krstic, S., Duma, A., Janevska, B., LEVKOV, Z., Nikolova, K. & Noveska, M. (2002): Diatoms in forensic expertise of drowning - a Macedonian experience. Forensic Science International 127(3): 198–203.
  5. LEVKOV, Z., Krstic, S., Nakov, T. & Melovski, Lj. (2005): Diatom assemblages on Shara and Nidze Mountains, Macedonia. Nova Hedwigia 81(3–4): 501–538.
  6. Karageorgis, A.P., Skourtos, M.S., Kapsamilis, V., Kontogianni, A.D., Skoulikidis, N.Th., Pagou, K., Nikolaidis N.P., Drakopoulou, P., Zanou, B., Karamanos, H., LEVKOV, Z. & Anagnostou, Ch. (2005): An integrated approach to watershed management within the DPSIR framefwork: Axios River catchemnt and Thermaikos Gulf. Regional Environmental Change 5: 138–160.
  7. LEVKOV, Z., Krstic, S. & Nakov, T. (2006): A new Gyrosigma species from Lakes Prespa and Ohrid, Republic of Macedonia. In: Ognjanova-Rumenova, N. & Manoylov, K. (eds): Fossil and Recent Phycological Studies pp. 139–145. Pensoft, Sofia.
  8. LEVKOV Z., Nakov, T & Metzeltin, D. (2006): New species and combination from genus Sellaphora Mereschkowsky (Bacillariophyceae) from Macedonia. Diatom Research 21: 297–312.
  9. LEVKOV, Z. & Williams, D.M. (2006): Two new species of Diatoma from Lakes Ohrid and Prespa, Macedonia. Diatom Research, 21: 281–296.
  10. Edlund, M.B., Brant, L.A., LEVKOV, Z. & Nakov, T. (2006): An emended description of Decussata (Patrick) Lange-Bertalot & Metzeltin that includes protoplast organization.Diatom Research 21: 297–310.
  11. LEVKOV, Z. Saul, B., Krstic, S., Nakov, T. & Ector, L. (2007): Ecology of benthic diatoms from Lake Prespa, Macedonia. Archiv für Hydrobiologie Supplement/ Algological Studies 124: 71–83.
  12. Krstic, S., Svircev, Z., LEVKOV, Z. & Nakov, T. (2007): Selecting appropriate bioindicator regarding the WFD guidelines for freshwaters – a Macedonian experience. International Journal on Algae 9: 41–63.
  13. LEVKOV, Z., Edlund, M.B. & Nakov, T. (2008): The identity and typification of Navicula hasta (Bacillariophyceae). Phycological Research, 56 (1): 46–57.
  14. LEVKOV, Z. & Nakov, T. (2008): Rhoicosphenia tenuis a new diatom species from Lake Ohrid. Diatom Research 23: 377–388.
  15. Nikolaidis, N.P., Karageorgis, A.P., Kapsimalis, V., Drakopoulou, P., Skoulikidis, N., Behrendt, H. & LEVKOV, Z. (2009): Management of nutrient emissions of Axios River catchment: Their effect in the coastal zone of Thermaikos Gulf, Greece. Ecological modeling 220: 383–396.
  16. Pavlov, A., Nakov, T., LEVKOV, Z., Lowe, R., Furey, P. & Ector, L. (2009): Luticola grupcei (Bacillariophyceae) - a new freshwater diatom from Mountain Baba (Macedonia) and Great Smoky Mountains National Park (USA): comparison with the type material of L. goeppertiana (Bleisch) D.G. Mann. Nova Hedwigia 89: 147–164.
  17. Edlund, M.B., Shinneman, A.L.C. & LEVKOV, Z. (2009): Diatom biodiversity in Mongolia: A new amphoroid diatom from saline lakes in western Mongolia, Amphora soninkhishigae sp. nov. Acta Botanica Croatica 68(2): 251–262.
  18. LEVKOV, Z. & Ector, L. (2010): A comparative study of Reimeria species (Bacillariophyceae). Nova Hedwigia 90(3–4): 469–489.
  19. LEVKOV, Z., Caput-Mihalic, K. & Ector, L. (2010): A taxonomical study of Rhoicosphenia Grunow (Bacillariophyceae) with a key for identification of taxa. Fottea 10: 1–56.
  20. Reed, J. M., Cvetkoska, A., LEVKOV, Z., Vogel, H. & Wagner, B. (2010): The last glacial-interglacial cycle in Lake Ohrid (Macedonia/Albania): testing diatom response to climate. Biogeosciences 7: 3083–3094.
  21. Ács, É., Ector, L. Kiss, K.T., Cserháti C., Morales, E.A. & LEVKOV, Z. (2011): Morphological observations and emended description of Amphora micrometra from the Bolivian Altiplano, South America. Diatom Research 26: 199–212.
  22. LEVKOV, Z. & Williams, D.M. (2011): Fifteen new diatom (Bacillariophyta) species from Lake Ohrid, Macedonia. Phytotaxa 30: 1–41.
  23. LEVKOV, Z. & Williams, D.M. (2012): Checklist of diatoms (Bacillariophyta) from Lake Ohrid and Lake Prespa (Macedonia), and their watersheds. Phytotaxa 45: 1–76.
  24. Pavlov, A. & LEVKOV, Z. (2013): Diversity and distribution of Eunotia Ehrenberg in Macedonia. Phytotaxa 86: 1–117.
  25. Krstić, S., Pavlov, A., LEVKOV, Z. & Jüttner, I. (2013): New Eunotia taxa in core samples from Lake Panch Pokhari in the Nepalese Himalaya. Diatom Research 28(2): 203–217.
  26. Pavlov, A., LEVKOV, Z., Williams, D.M. & Edlund, M.B. (2013): Observations on Hippodonta (Bacillariophyceae) in selected ancient lakes. Phytotaxa 90: 1–53.
  27. Jovanovska, E., Nakov, T. & LEVKOV, Z. (2013): Observations of the genus Diploneis (Ehrenberg) Cleve from Lake Ohrid, Macedonia. Diatom Research 28(3): 237-262.
  28. Jovanovska, E., Buczkó, K., Ognjanova-Rumenova, N., Nakov, T. & LEVKOV, Z. (2013): Identity and typification of Diploneis ostracodarum, Diploneis budayana and Diploneis praeclara (Bacillariophyta). Phytotaxa 137: 15–26.
  29. Pavlov A. & LEVKOV, Z. (2013): Observations on the genus Pinnularia section Distantes (Bacillariophyta) from Macedonia; diversity and distribution. Contributions, Section of Natural, Mathematical and Biotechnical Sciences, MASA Vol. 34: 33–57.
  30. Cvetkoska, A., LEVKOV, Z. & Hamilton, P.B. (2014): Surirella subrotunda sp. nov. and Surirella parahelvetica sp. nov., two new diatom (Bacillariophyta) species from Lake Prespa, Macedonia. Phytotaxa 156: 145–155.
  31. Tofilovska, S., Wetzel, C.E., Ector, L. & LEVKOV, Z. (2014): Observation on Achnanthes Bory sensu stricto (Bacillariophyceae) from subaerial habitats in Macedonia and comparison with the type material of A. coarctata (Brébisson ex W. Smith) Grunow, A. coarctata var. sinaensis Hustedt and A. intermedia Kützing. Fottea 14(1): 15–42.
  32. Cvetkoska, A., Hamilton, P.B., Ognjanova–Rumenova, N. & LEVKOV, Z. (2014): Observations of the genus Cyclotella (Kützing) Brébisson in ancient lakes Ohrid and Prespa and a description of two new species C. paraocellata sp. nov. and C. prespanensis sp. nov. Nova Hedwigia 98 (3–4): 313–340.
  33. Wagner, B., Wilke, T., Krastel, S., Zanchetta, G., Sulpizio, R. Reicherter, R., Leng, M. J., Grazhdani, A., Trajanovski, S., Francke, A., Lindhorst, K., LEVKOV, Z., Cvetkoska, A., Reed, J. M., Zhang, X., Lacey, J. H., Wonik, T., Baumgarten, H. & Vogel, H. (2014): The SCOPSCO drilling project recovers more than 1.2 million years of history from Lake Ohrid. Scientific Drilling 17: 19–29.
  34. LEVKOV, Z. & Williams, D.M. (2014): Observations on Caloneis Cleve (Bacillariophyceae) species from the ancient lakes Ohrid and Prespa. Nova Hedwigia Beiheft 143: 141–158.
  35. Cvetkoska, A., LEVKOV, Z., Reed, J.M. & Wagner, B. (2014): Late glacial to Holocene climate change and human impact in the Mediterranean: the last ca. 17 ka diatom record of Lake Prespa (Macedonia/Albania/Greece). Palaeogeography, Palaeoclimatology, Palaeoecology 406: 22–32.
  36. Zidarova, R., LEVKOV, Z. & Van de Vijver, B. (2014): Four new Luticola taxa (Bacillariophyta) from Maritime Antarctica. Phytotaxa 170(3): 155–168.
  37. Voigt, R. & LEVKOV, Z. (2014): Halamphora saxonica sp. nov. (Bacillariophyceae) from lakes in lignite-mining area in Saxony, Germany. Nova Hedwigia 99(1–2): 213–222.
  38. Mitić-Kopanja, D., Wetzel, C.E., Ector, L. & LEVKOV, Z. (2014): Two new Gomphonema Ehrenberg (Bacillariophyceae) species from Macedonia and comparison with type material of G. brebissonii Kützing. Fottea 14(2): 149–160.
  39. Van de Vijver, B., Kopalová, K., Zidarova, R. & LEVKOV, Z. (2014): Revision of the genus Halamphora (Bacillariophyta) in the Antarctic Region. Plant Ecology and Evolution 147(3): 374–391
  40. Zhang, X., Reed, J. Wagner, B., Francke, A. & LEVKOV, Z. (2014): Lateglacial and Holocene climate and environmental change in the northeastern Mediterranean region: diatom evidence from Lake Dojran (Republic of Macedonia/Greece). Quaternary Science Reviews 103: 51–66.
  41. Cvetkoska, A., LEVKOV, Z., Hamilton, P.B. & Potapova, M. (2014): The biogeographic distribution of Cavinula (Bacillariophyceae) in North America with the descriptions of two new species. Phytotaxa 184(4): 181–207.
  42. Peng, Y. Rioual, P., LEVKOV, Z., Williams, D.M. & Jin, Z. (2014): Morphology and ultrastructure of Hippodonta qinghainensis sp. nov. (Bacillariophyceae), a new diatom from Lake Qinghai, China. Phytotaxa 186(2): 61–74.
  43. Jovanovska, E., Buczkó, K., Nakov, T. & LEVKOV, Z. (2014): Diploneis transylvanica (Bacillariophyceae), a new diatom species from the Neogene fossil deposit in Romania. Nova Hedwigia 99(3–4): 413–427.
  44. Cvetkoska, A., LEVKOV, Z., Reed, J.M., Wagner B., Panagiotopoulos, K., Leng, M.J. & Lacey, J.H. (2015): Quaternary climate change and Heinrich Events in the southern Balkans: Lake Prespa diatom palaeolimnology from the last interglacial to present. Journal of Paleolimnology 53: 215–231.
  45. Ognjanova-Rumenova, N., Jovanovska, E., Cvetkoska, A. & LEVKOV, Z. (2015): Two new Tertiarius (Bacillariophyta, Coscinodiscophyceae) species from Mariovo Neogene Basin, Macedonia. Fottea 15(1): 51–62.
  46. Jovanovska, E., & LEVKOV, Z., & Edlund, M.B. (2015): The genus Diploneis Ehrenberg ex Cleve (Bacillariophyta) from Lake Hövsgöl, Mongolia. Phytotaxa 217(3): 201–248.
  47. Cvetkoska, A., Hamilton, P.B., & LEVKOV, Z. (2015): Surirella prespanensis sp. nov. and Surirella hinziae sp. nov., two new diatom (Bacillariophyceae) species from ancient Lake Prespa (Macedonia/Albania/Greece). Fottea 15(2): 179–189.
  48. Jüttner, I., Williams, D.M., LEVKOV, Z., Falasco, E., Battegazzore, M., Cantonati, M., Van de Vijver, B., Angele, C. & Ector, L. (2015): Reinvestigation of the type material for Odontidium hyemale (Roth) Kützing and related species, with description of four new species in the genus Odontidium (Fragilariaceae, Bacillariophyta). Phytotaxa 234(1): 1–36.
  49. Jovanovska, E., Cvetkoska, A., Tofilovska, S., Ognjanova- Rumenova & LEVKOV, Z. (2016): Description of a new fossil diatom genus, Cribrionella gen. nov. (Bacillariophyta) from Quaternary sediments of Lake Ohrid. Phytotaxa 252(1): 31–42.
  50. Jovanovska, E., Cvetkoska, A., Hauffe, T., LEVKOV, Z., Wagner, B., Sulpizio, R., Francke, A., Albrecht, C. & Wilke, T. (2016): Differential resilience of ancient sister lakes Ohrid and Prespa to environmental disturbances during the Late Pleistocene. Biogeosciences 13: 1149–1161.
  51. Zhang, X.S., Reed, J.M., Lacey, J.H., Francke, A., Leng, M.J., LEVKOV, Z. & Wagner, B. (2016): Complexity of diatom response to Lateglacial and Holocene climate and environmental change in ancient, deep and oligotrophic Lake Ohrid (Macedonia and Albania). Biogeosciences 13: 1351–1365.
  52. Pavlov, A., Jovanovska, E., Wetzel, C.E., Ector, L. & LEVKOV, Z. (2016): Freshwater Mastogloia (Bacillariophyceae) taxa from Macedonia, with a description of the epizoic M. sterijovskii sp. nov. Diatom Research 31(2): 85–112.
  53. Cvetkoska, A., Jovanovska, E., Francke, A., Tofilovska, S., Vogel, H., LEVKOV, Z., Donders, T.H., Wagner, B. & Wagner-Cremer, F. (2016): Ecosystem regimes and responses in a coupled ancient lake system from MIS 5b to present: the diatom record of lakes Ohrid and Prespa. Biogeosciences 13: 3147–3162.
  54. Tofilovska, S., Cvetkoska, A., Jovanovska E., Ognjanova-Rumenova, N. & LEVKOV, Z. (2016): Two new fossil Cyclotella (Kützing) Brébisson species from Lake Ohrid, Macedonia/Albania. Fottea 16(2): 218–233.
  55. LEVKOV, Z., Tofilovska, S., Jovanovska, E., Cvetkoska, A. & Metzeltin, D. (2016): Revision of the Stauroneis smithii Grunow (Bacillariophyceae) species complex from Macedonia. Botanica Serbica 40(2): 167–178.
  56. Wilke, T., Wagner, B., Van Bocxlaer, B., Albrecht, C., Ariztegui, D., Delicado, D., Francke, A., Harzhauser, M., Hauffe, T., Holtvoeth, J., Just, J., Leng, M.J., LEVKOV, Z., Penkman, K., Sadori, L., Skinner, A., Stelbrink, B., Vogel, H., Wesselingh, F. & Wonik, T. (2016): Scientific drilling projects in ancient lakes: integrating geological and biological histories. Global and Planetary Change 143: 118–151.
  57. LEVKOV, Z., Tofilovska, S., & Mitić-Kopanja, D. (2016): The diatom genus Craticula Grunow (Bacillariophyceae) from Macedonia. Contributions, Section of Natural, Mathematical and Biotechnical Sciences 37(2): 129–165.
  58. Kale, A., LEVKOV, Z. & Karthick, B. (2017): Typification of two species of Luticola (Bacillariophyta) from aerophilic habitats of the Western Ghats, India. Phytotaxa 298(1): 29–42.
  59. Jüttner, I, Williams, D.M., Gurung, S., Van de Vijver, B., LEVKOV, Z., Sharma, C.M., Shara, S., Cox, E.J. (2017): The genus Odontidium (Bacillariophyta) in the Himalaya - a preliminary account of some taxa and their distribution. Phytotaxa 332: 1–21.
  60. Wagner, B., Wilke, T., Francke, A., Albrecht, C., Baumgarten, H., Bertini, A., Combourieu-Nebout, N., Cvetkoska, A., D'Addabbo, M., Donders, T.H., Föller, K., Giaccio, B., Grazhdani, A., Hauffe, T., Holtvoeth, J., Joannin, S., Jovanovska, E., Just, J., Kouli, K., Koutsodendris, A., Krastel, S., Lacey, J. H., Leicher, N. Leng, M.J., LEVKOV, Z., Lindhorst, K., Masi, S., Mercuri, A.M., Nomade, S., Nowaczyk, N., Panagiotopoulos, K., Peyron, O., Reed, J.M. Regattieri, E., Sadori, L., Sagnotti, L., Stelbrink, B., Sulpizio, R., Tofilovska, T., Torri, P., Vogel, H., Wagner, T., Wagner-Cremer, F., Wolff, G. A., Wonik, T., Zanchetta, G. & Zhang X. S. (2017): The environmental and evolutionary history of Lake Ohrid (FYROM/Albania): interim results from the SCOPSCO deep drilling project. Biogeosciences 14: 2033–2054.
  61. LEVKOV, Z., Williams, D.M., Nikolovska, D, Tofilovska, S. & Čakar, Z. (2017): The use of diatoms in forensic science: Advantages and limitations of the diatom test in cases of drowning. In: Williams, M., Hill, T., Boomer, I. & Wilkinson, I. (Eds): The Archaeological and Forensic Applications of Microfossils: A Deeper Understanding of Human History. The Micropalaeontological Society Special Publications. Geological Society, London, 261–277.
  62. LEVKOV, Z., Tofilovska, S., Wetzel, C.E., Mitić-Kopanja, D., & Ector, L. (2017): Diversity of Luticola D.G. Mann (Bacillariophyceae) species on halomorphic soils from Gladno Pole, Central Macedonia. Nova Hedwigia Beiheft 146: 175–196.
  63. LEVKOV, Z. & Williams D.M. (2017): Caloneis parva Levkov & D.M.Williams, nom. nov., a new name for Caloneis minuta Levkov & D.M.Williams. Notulae algarum 39: 1.
  64. Cvetkoska, A., Pavlov, A., Jovanovska, E., Tofilovska, S., Blanco, S., Ector, L., Wagner-Cremer, F. & LEVKOV, Z. (2018): Spatial patterns of diatom diversity and community structure in ancient Lake Ohrid. Hydrobiologia 819: 197–215.
  65. Stelbrink, B., Jovanovska, E., LEVKOV, Z., Ognjanova-Rumenova, N., Wilke, T. & Albrecht, C. (2018): Diatoms do radiate: evidence for a freshwater species flock. Journal of Evolutionary Biology 31: 1969–1975.
  66. Jüttner, I., Kociolek, J.P., Gurung, S., Gurung, A., Mani Sharma, C., LEVKOV, Z., Williams, D.M. & Ector, L. (2018): The genus Gomphonema (Bacillariophyta) in Rara Lake, Nepal: taxonomy, morphology, habitat distribution and description of five new species. Diatom Research 33: 1–38.
  67. Van de Vijver, B., Wetzel, C.E., LEVKOV, Z. & Ector, L. (2019): Analysis of the type slide of Achnanthes boyei Østrup (Achnanthidiaceae, Bacillariophyta). Notulae Algarum 96: 1–6.
  68. Caput Mihalić, K., Gligora Udovič, M., Galović, I., Stanković, I., Šušnjara, M., Žutinić, P., Kulaš, A., Špoljarić, I. & LEVKOV, Z. (2019): Tetramphora croatica sp. nov. – A new brackish-water species from Lake Vransko, Croatia. Phytotaxa 401: 276–286.
  69. Vidakovic, D., Hamilton, P.H. & LEVKOV, Z. (2019): Neidiopsis borealis sp. nov, a new diatom species from the mountain Shar Planina, Macedonia. Phytotaxa 402: 21–28.
  70. Zhang, W., Wang, T., LEVKOV, Z., Jüttner, I., Ector, L. & Zhou, Q-C (2019): Halamphora daochengensis sp. nov., a new freshwater diatom species (Bacillariophyceae) from a small mountain lake, Sichuan Province, China. Phytotaxa 404: 012–022.
  71. LEVKOV, Z., Vidaković, D., Cvetkoska, A., Mitić-Kopanja, D., Krstić, S., Van de Vijver, B. & Hamilton, P. B. (2019): Observations of the genus Muelleria (Frenguelli) Frenguelli (Bacillariophyceae) from the Republic of North Macedonia. Plant ecology and Evolution 152(2): 293–312.
  72. Cantonati, M., Angeli, N., Lange-Bertalot, H. & LEVKOV, Z. (2019): New Amphora and Halamphora species from springs in the northern Apennines (Emilia-Romagna, Italy). Plant ecology and Evolution 152(2): 285–292.
  73. Kulichová, J., Neustupa, J., Vrbová, K., LEVKOV, Z. & Kopalová, K. (2019): Asymmetry in Luticola species. Diatom Research 34(2): 67–74.
  74. Wagner, B., Vogel, H., Francke, A., Friedrich, T., Donders, T., Lacey, J.H., Leng, M.J., Regattieri, E., Sadori, L., Wilke, T., Zanchetta, G., Albrecht, C., Bertini A., Combourieu-Nebout, N., Cvetkoska, A., Giaccio, B., Grazhdani G., Hauffe, T., Holtvoeth, J., Joannin, S., Jovanovska E., Just, J. Kouli, K., Kousis, I., Koutsodendris, A., Krastel, S., Lagos, M., Leicher, N., LEVKOV, Z., Lindhorst, K., Masi, A., Melles, M.,. Mercuri, A.M., Nomade, S., Nowaczyk, N., Panagiotopoulos, K., Peyron, O., Reed, J.M., Sagnotti, L., Sinopoli, G. Stelbrink, B., Sulpizio, R., Timmermann, A., Tofilovska, S., Torri, P., Wagner-Cremer, F., Wonik T. & Zhang, X. (2019): Mediterranean winter rainfall in phase with African monsoons during the past 1.36 million years. Nature 573: 256–260.
  75. Cantonati, M., Poikane, S., Pringle, C.M., Stevens, E.L., Turak, E., Heino, J., Richardson, J., Bolpagni, R. Borrini, A., Cid, N., Čtvrtlíková, M., Galassi, D.M.P., Hájek, M., Hawes, I., LEVKOV, Z., Naselli-Flores, L. Saber, A.A., Di Cicco, M., Fiasca B., Hamilton, P.B., Kubečka, J. Segadelli, S. Znachor, P. (2020): Multiplicity, Characteristics, Main Impacts, and Stewardship of Natural and Artificial Freshwater Environments: Consequences for Biodiversity Conservation. Water 12: 1–82.
  76. Wilke, T., Hauffe, T, Jovanovska, J., Cvetkoska, A., Donders, T., Ekschmitt, K., Francke, A., Lacey.J.H., LEVKOV, Z., Marshall, C.R., Neubauer. T.A., Silvestro, D., Stelbrink, B., Vogel, H., Albrecht, C., Holtvoeth, J., Krastel, S., Leicher, N., Leng, M.J., Lindhorst, K., Masi, A., Ognjanova-Rumenova, N., Panagiotopoulos, K., Reed, J.M., Sadori, L., Tofilovska, S., Van Bocxlaer, B., Wagner-Cremer, F., Wesselingh, F.P., Wolters, V., Zanchetta, G., Zhang, X. & Wagner, B. (2020): Deep drilling reveals massive shifts in evolutionary dynamics after formation of ancient ecosystem. Science Advances 6 : eabb2943.
  77. Caput Mihalić, K., Galović, I., Wetzel, C.E., Ector, L., Ilijanić, N., Miko, S., Wishkerman, A., Hamilton, P.B. & LEVKOV, Z. (2020): Envekadea vranaensis sp. nov. a new diatom species (Bacillariophyta) from the lacustrine Holocene sediments of Lake Vrana, Croatia. Nova Hedwigia 110(1–2): 1–19.
  78. Fernádez-Moreno, D., Luís, A.T. Sánchez-Castillo, P.M., Vidakovic, D. & LEVKOV, Z. (2019): Pinnularia baetica sp. nov. (Bacillarophyceae): revision of similar Pinnularia genus taxa and Mediterranean distribution of a panduriform diatom. Phytotaxa 435(2): 085–100.
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  81. Schabetsberger, R. Grapci-Kotori, L., Ibrahimi, H., Bilalli, A., LEVKOV, Z., Jersabek, C.D., Vorage, M., Denoël, M., Kaiser, R., Pall, K., Eisendle, U., Eder, E. & Sadiku, A. (2021): First limnological characterization of Lakes Leqinat and Drelaj in Bjeshkët e Nemuna National Park, Kosovo. Eco.Mont. 13: 12–21.
  82. Schabetsberger, R., Jersabek, C.D., LEVKOV, Z., Ehrenfellner, B., Enoka, L.F. & Faiilagi, S.A. (2021): Limnological Characterization of Three Tropical Crater Lakes in the Archipelago of Samoa (Lanoto'o, Olomaga, Mataulano). Pacific Science 75(1), 163-174.
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  87. Gligora Udovic, M., Kulas, A., Susnjara, M., Arapov, J., Blanco. S., & LEVKOV, Z. (2022): Cymbopleura amicula stat nov. et. nom. nov. (Bacillariophyceae)-a rare diatom species from a karst river in Croatia. Phytotaxa 532 (2): 139–151.
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  90. Zaova, D., Cvetkoska, A., Wagner, B., Francke, A., Vogel, H., LEVKOV, Z., Jovanovska, E. (2022): Diatom community responses to environmental change in Lake Ohrid (Balkan Peninsula) during the mid-Pleistocene Transition. Quaternary International 622: 1–9.
  91. Stankovic, I., Gligora Udovic, M., Susnjara, M., Alegro, A. & LEVKOV, Z. (2022): Diversity and ecology of benthic diatoms in a mire with narrow pH gradient. Fottea 22(1): 39–55.
  92. Vidaković, D., Krizmanić, J., Gavrilović, B., Dojčinović, B.P., LEVKOV, Z., Dimkić I. & Ćirić, M. (2022): Artificial sandpit lake as a habitat of brackish diatom species. Botany Letters 169(3): 360–369.
  93. Videska, A., Zaova, D., Naumovska, H., Kochoska, H., Mitic-Kopanja, D. & LEVKOV, Z. (2022): Cymbella biseriata sp. nov.—a unique living species of Cymbella Agardh (Bacillariophyceae) with biseriate striae. Phytotaxa 561(1): 41–52.
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